In the multiverse called the data world, everything corresponding to the real world is stored. Within it, the data from relics of human culture gained self-awareness, and a secret group of spirits established an alliance: The Country of Rare Treasure.

Upon hearing news that islands in East Blue are being destroyed, Monkey D. Luffy and his crew go to investigate. On their way, however, an outlandish pirate ship appears out of the sky, helmed by the infamous pirate Shiki "the Golden Lion"—a man who ate the Float-Float Fruit and the first ever prisoner to escape from Impel Down. In his quest to defeat the World Government, Shiki kidnaps Nami to be his own navigator and sends the rest of the Straw Hat Pirates to his floating islands as hostages, leaving her in a dilemma. Separated in a land under Shiki's absolute control, Luffy and his crew must survive the mystifying terrain in order to bring back their navigator and friend.

The world's greatest exposition of the pirates, by the pirates, for the pirates—the Pirates Festival. Luffy and the rest of the Straw Hat Crew receive an invitation from its host Buena Festa who is known as the Master of Festivities. They arrive to find a venue packed with glamorous pavilions and many pirates including the ones from the Worst Generation. The place is electric.

High above the realm of man, the gods of the world have convened to decide on a single matter: the continued existence of mankind. Under the head of Zeus, the deities of Ancient Greece, Norse mythology, and Hinduism, among others, call assembly every one thousand years to decide the fate of humanity. Because of their unrelenting abuse toward each other and the planet, this time the gods vote unanimously in favor of ending the human race. But before the mandate passes, Brunhild, one of the 13 demigod Valkyries, puts forth an alternate proposal: rather than anticlimactically annihilating mankind, why not give them a fighting chance and enact Ragnarök, a one-on-one showdown between man and god? Spurred on by the audacity of the challenge, the divine council quickly accepts, fully confident that this contest will display the utter might of the gods. To stand a chance against the mighty heavens, Brunhild will need to assemble history's greatest individuals, otherwise the death knell will surely be sounded for mankind.